Gramma, the beloved Galapagos tortoise who lived at the San Diego Zoo for nearly a century, has died on Nov. 20. She was (about) 141. The zoo's care team "made the compassionate and exceptionally ...

Gramma, a 141-year-old Galapagos tortoise and the San Diego Zoo’s oldest resident, has died. She came from the Bronx Zoo nearly a century ago and was known as the "Queen of the Zoo." Visitors and ...

The San Diego Zoo has said goodbye to one of its most cherished icons, Gramma, a Galápagos tortoise who inspired generations of visitors and conservationists. She passed away on November 20 after care ...

(KSWB/KUSI) — Gramma, a Galapagos tortoise that was the oldest living animal at the San Diego Zoo, has died. The zoo said Gramma had ongoing bone conditions and that caretakers had to make the ...

SAN DIEGO — One of the first animals brought to the San Diego Zoo has died at the age of 141 years old. The zoo announced the death of its beloved Galápagos tortoise, Gramma, on social media, saying ...

LOS ANGELES — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.